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What do these two fields do? I already have the option to select WHO to send the reply to and FROM which email it comes from. What is the purpose for this then

Hi @raghavbajoria123 

The From Name is the label for the From Email.

e.g. Zapier Support for support@zapier.com

 

The Reply To would be a different email address than the email address that send the email.

If a recipient replies the email would be sent to the Reply To email address.